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Osteogenic markers, including BMP-2, Osterix, Collagen type I and Osteocalcin, are upregulated to induce osteogenesis (82,83)

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During the lysis of autophagosomes, ROC-325 can inhibit the formation of autophagolysosomes by increasing lysosomal membrane permeability and lysosomal deacidification, and it can significantly reduce the autophagic flux in PAECs and activate the nitric oxide signaling pathway, which in turn improves right ventricular dysfunction and pulmonary vascular remodeling in hypoxic PH rats [100]